1. National Museum of Marine Biology & Aquarium; No. 2, Houwan Road Checheng Pingtung 94450 Taiwan
2. Division of Biodiversity & Evolution; Institute of Marine Biology; National Dong Hwa University; Checheng Pingtung 94450 Taiwan
3. Aquatic Ecology Laboratory; The Ohio State University; 1314 Kinnear Road Columbus OH 43212-1156 USA