1. Key Laboratory of Aquatic Biodiversity and Conservation of Chinese Academy of Sciences; Institute of Hydrobiology; Chinese Academy of Sciences; Wuhan 430072 China
2. Institute of Evolution & Marine Biodiversity; Ocean University of China; Qingdao 266003 China
3. School of Life Sciences; East China Normal University; Shanghai 200241 China
4. Department of Life Sciences; Natural History Museum; Cromwell Road London SW7 5BD United Kingdom