1. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology Yunnan University China
2. State Key Laboratory for Conservation and Utilization of Bioresources in Yunnan Yunnan University China
3. Department of Evolution and Ecology University of California Davis Davis CA 95616 USA
4. Department of Biology Stanford University Stanford CA 94305 USA
5. Department of Genetics University of North Carolina Chapel Hill NC 27599 USA
6. The Hokkaido University Museum Kita‐10, Nishi‐8 Kitaku, Sapporo 060–0810 Japan