1. School of Biosciences; University of Nottingham; Sutton Bonington Nottinghamshire LE12 5RD UK
2. Wageningen University and ISRIC - World Soil Information, Droevendaalsesteeg 3; 6708 PB Wageningen the Netherlands
3. School of Life and Environmental Sciences; Sydney Institute of Agriculture, The University of Sydney; New South Wales 2006 Australia