1. CAS Key Laboratory for Plant Biodiversity and Biogeography of East Asia Kunming Institute of Botany, Chinese Academy of Sciences Kunming Yunnan 650201 China
2. Ministry of Education Key Laboratory for Transboundary Ecosecurity of Southwest China and Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology Institute of Biodiversity, School of Ecology and Environmental Science, Yunnan University Kunming Yunnan 650504 China
3. Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations Kunming Institute of Botany, Chinese Academy of Sciences Kunming Yunnan 650201 China
4. College of Wildlife and Protected Area Northeast Forestry University Harbin Heilongjiang 150040 China
5. Department of Biosciences Durham University South Road Durham DH1 3LE UK