1. Exotic Disease Inspection Division; Laboratory Department; Animal Quarantine Service; Ministry of Agriculture, Forestry and Fisheries; Tokoname Aichi Japan
2. Laboratory of Microbiology; Department of Disease Control; Graduate School of Veterinary Medicine; Hokkaido University; Sapporo Hokkaido Japan
3. Microbiological Examination Division; Laboratory Department; Animal Quarantine Service; Ministry of Agriculture, Forestry and Fisheries; Yokohama Kanagawa Japan
4. Global Station for Zoonosis Control; Global Institution for Collaborative Research and Education (GI-CoRE); Hokkaido University; Sapporo Hokkaido Japan