1. Department of Conservation Ecology and Entomology; Stellenbosch University; Matieland South Africa
2. South African National Biodiversity Institute; Kirstenbosch Research Centre; Claremont South Africa
3. Percy FitzPatrick Institute of African Ornithology; DST/NRF Centre of Excellence; Department of Biological Sciences; University of Cape Town; Rondebosch South Africa
4. CNRS; Sorbonne Universités; UPMC Univ Paris 06; Institute of Ecology and Environmental Sciences; Paris France