1. Life Science Center of Tsukuba Advanced Research Alliance (TARA Center); University of Tsukuba; Tsukuba Ibaraki 305-8577 Japan
2. Developmental Genetics; National Institute for Basic Biology; Higashiyama Okazaki 444-8787 Japan
3. Graduate School of Life and Environmental Sciences; University of Tsukuba; Tsukuba Ibaraki 305-8572 Japan