1. State Key Laboratory of Plant Physiology and Biochemistry; College of Life Sciences; Zhejiang University; Hangzhou 310058 China
2. State Key Laboratory of Rice Biology; China National Rice Research Institute; Chinese Academy of Agricultural Sciences; Hangzhou 310006 China
3. Department of Biology - Plant Biology; University of Fribourg; Rue Albert-Gockel 3 Fribourg CH-1700 Switzerland