1. Department of Environmental Science, Policy, and Management University of California Berkeley Berkeley CA USA
2. Museum of Vertebrate Zoology University of California Berkeley Berkeley CA USA
3. Department of Biological Sciences University of Pittsburgh Pittsburgh PA USA
4. Department of Biology University of Nevada Reno Reno NV USA
5. Smithsonian Tropical Research Institute Panamá República de Panamá
6. Sistema Nacional de InvestigaciónSENACYT Clayton Panamá República de Panamá