1. Université de Toulouse; INP, UPS, EcoLab (Laboratoire Ecologie Fonctionnelle et Environnement); Toulouse France
2. CNRS, EcoLab; Toulouse France
3. MARE - Marine and Environmental Sciences Centre; Department of Life Sciences; University of Coimbra; Coimbra Portugal
4. Centre for Functional Ecology; Department of Life Sciences; University of Coimbra; Coimbra Portugal