1. Department of Ecological Sciences Section Animal Ecology Vrije Universiteit Amsterdam De Boelelaan 1085 Amsterdam 1081 HV The Netherlands
2. Institute for Evolution and Biodiversity University of Münster Hüfferstraße 1 Münster 48149 Germany
3. Bioinformatics University of Applied Sciences Leiden Zernikedreef 11 Leiden 2333 CK The Netherlands