1. INRA; UMR 1355; Sophia Antipolis F-06903 France
2. Université de Nice Sophia Antipolis; UMR Institut Sophia Agrobiotech; Sophia Antipolis France
3. CNRS; UMR 7254; Sophia Antipolis France
4. CIRAD; UMR BGPI; Campus International de Baillarguet; Montpellier F-34398 France
5. University College London; London WC1E 6BT UK