1. Graduate School of Frontier Biosciences Osaka University 1‐3 YamadaokaSuita Osaka 565‐0871Japan
2. Division of Biological Science, Graduate School of Science Nagoya University Chikusa‐kuNagoya 464‐8602Japan
3. RIKEN Center for Biosystems Dynamics Research & SPring‐8 Center 1‐3 YamadaokaSuita Osaka 565‐0871Japan