1. Key Laboratory of Sustainable Forest Ecosystem Management, Center for Ecological Research, Ministry of Education Northeast Forestry University Harbin China
2. CAS Key Laboratory of Forest Ecology and Management Institute of Applied Ecology, Chinese Academy of Sciences Shenyang China
3. Agronomy Department University of Florida Gainesville Florida USA
4. Department of Ecology & Evolutionary Biology High Meadows Environmental Institute, Princeton University Princeton New Jersey USA
5. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems and Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Chinese Academy of Sciences Guangzhou China