1. Institute of Zoology, University of Bern, Baltzerstrasse 6, 3012 Bern, Switzerland;
2. Department of Ecology & Evolution, University of Lausanne, Biology Building, 1015 Lausanne, Switzerland;
3. Department of Applied Biology, Estación Biológica de Doñana, Pabellón del Perú, Avda. M. Luisa s/n, E‐41013 Sevilla, Spain
4. Institute of Zoology, University of Neuchâtel, Rue Emile‐Argand 11, 2007 Neuchâtel, Switzerland