1. Institute of Hydrobiology; Chinese Academy of Sciences; 7 South Donghu Road, Wuchang District Wuhan 430072 China
2. University of Chinese Academy of Sciences; 19 Yuquan Road, Shijingshan District Beijing 100049 China
3. School of Life Sciences, Anqing Normal University; 1318 North Jixian Road, Yixiu District Anqing 246133 China
4. Marine Biology Institute, College of Science; Shantou University; 243 Daxue Road, Shantou 515063 China