1. School of Agriculture and Environment and the UWA Institute of Agriculture; Faculty of Science; The University of Western Australia; 35 Stirling Highway Crawley WA 6009 Australia
2. Plant Biosecurity Cooperative Research Centre (PBCRC); Bruce ACT 2617 Australia
3. School of Veterinary and Life Sciences; Murdoch University; South Street Murdoch WA 6105 Australia
4. Department of Agriculture and Food Western Australia; 3 Baron-Hay Court South Perth WA 6151 Australia