1. Key Laboratory of Plant Resources and Beijing Botanical Garden; Institute of Botany; Chinese Academy of Sciences; Beijing 100093 China
2. University of Chinese Academy of Sciences; Beijing 100049 China
3. State Key Laboratory of Systematic and Evolutionary Botany; Institute of Botany; Chinese Academy of Sciences; Beijing 100093 China
4. Key Laboratory of Plant Germplasm Enhancement and Speciality Agriculture; Wuhan Botanical Garden; Chinese Academy of Sciences; Wuhan 430074 China