1. Ecology and Evolutionary Biology, Yale Univ. New Haven CT USA
2. Center for Biodiversity and Global Change, Yale Univ. New Haven CT USA
3. Ecology and Evolutionary Biology, Univ. of Connecticut Storrs CT USA
4. School of Forestry and Environmental Studies, Yale Univ. New Haven Connecticut USA
5. Vertebrate Ecology and Evolution Research Group, Inst. of Biology, Univ. de Antioquia Medellín Colombia