1. School of Biological Sciences University of Bristol Life Sciences Building 24 Tyndall Avenue Bristol BS8 1TQ UK
2. Institute of Systematics, Evolution and Biodiversity CNRS Natural History Museum Paris UPMC Sorbonne University EPHE 57 rue Cuvier 75005 Paris France
3. Department of Plant Sciences University of Cambridge Downing Street Cambridge CB2 3EA UK
4. Royal Botanic Garden Edinburgh 20a Inverleith Row Edinburgh EH3 5LR UK
5. UMR 3330 UMIFRE 21 French Institute of Pondicherry CNRS 11 Saint Louis Street Pondicherry 605001 India