1. Tanzania Wildlife Research Institute; P.O Box 661 Arusha Tanzania
2. Department of Integrative Biology; University of Guelph; 50 Stone Road East Guelph Ontario N1G 2W1 Canada
3. Wilderness & Wildlife Conservation Trust; No.130, Reid Avenue Colombo Srilanka
4. Department of Zoology and Wildlife Conservation; University of Dar es Salaam; P.O Box 35064 Dar es Salaam Tanzania
5. Center for Biodiversity Research; University of British Columbia; 6270 University Boulevard Vancouver British Columbia V6TIZ4 Canada