1. Laboratoire MIVEGEC (UMR CNRS 5290, UR IRD 224, UM1, UM2); 911 avenue Agropolis, BP 64501 Montpellier Cedex 5 34394 France
2. Department of Zoology; University of Otago; 360 Great King Street Dunedin New Zealand
3. Molecular Ecology and Evolution Lab; Department of Biology; Lund University; SE-223 62 Lund Sweden
4. The Charles Perkins Center; The University of Sydney; Sydney NSW 2006 Australia
5. School of Biological Sciences; The University of Sydney; Sydney NSW 2006 Australia