1. INRA and Université de Lorraine; UMR Interactions Arbres/Micro-organismes 1136; F-54280 Champenoux France
2. Clermont Université; Université Blaise-Pascal; UMR 547 PIAF, BP 10448, F-63000 Clermont-Ferrand France
3. INRA; UMR 547 PIAF, F-63100 Clermont-Ferrand France
4. Umeå Plant Science Center, Department of Forest Genetics and Plant Physiology; The Swedish University of Agricultural Sciences; SE-901 83 Umeå Sweden