1. Department of Biology McGill University Montreal Quebec H3A 1B1 Canada
2. Smithsonian Tropical Research Institute Apartado 0843–03092 Balboa Ancón Republic of Panama
3. Current Address: GEOMAR Helmholtz Centre for Ocean Research Kiel Evolutionary Ecology of Marine Fishes Düsternbrooker Weg 20 Kiel 24105 Germany
4. Current Address: Museum of Natural Science, Department of Biological Sciences Louisiana State University 119 Foster Hall Baton Rouge LA
5. Current Address: Patricia and Phillip Frost Museum of Science 3280 South Miami Avenue Miami FL