1. Jiangsu Co‐Innovation Center for Modern Production Technology of Grain Crops Yangzhou University 225009 Yangzhou China
2. State Key Lab of Plant Genomics, Institute of Genetics and Developmental Biology, Innovation Academy for Seed Design Chinese Academy of Sciences 100101 Beijing China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. Institute of Food Crops Hubei Academy of Agricultural Sciences Wuhan 430064 China
5. Key Laboratory of Cell Proliferation and Regulation Biology of Ministry of Education, College of Life Science Beijing Normal University Beijing 100875 China