1. Arnold Arboretum of Harvard University Boston MA 02131 USA
2. Department of Organismic and Evolutionary Biology Harvard University Cambridge MA 02138 USA
3. Global Change Ecology and Evolution (GloCEE) Department of Life Sciences University of Alcalà Alcalà de Henares 28805 Spain
4. Forest & Conservation Sciences Faculty of Forestry University of British Columbia Vancouver BC V6T 1Z4 Canada