1. School of Earth, Environmental and Biological Sciences; Queensland University of Technology; Brisbane QLD 4001 Australia
2. Plant Biosecurity Cooperative Research Centre; Bruce ACT Australia
3. Centre for Microscopy and Microanalysis; The University of Queensland; Brisbane QLD 4072 Australia
4. School of Biological Sciences; The University of Queensland; Brisbane QLD 4072 Australia
5. Institute for Future Environments; Queensland University of Technology; Brisbane QLD 4001 Australia
6. Department of Agriculture, Fisheries and Forestry, Ecosciences Precinct; GPO Box 267 Brisbane QLD 4001 Australia