1. CNRS; Laboratoire d'Ecologie Alpine (LECA); UMR-CNRS 5553; Université J. Fourier; BP 53 38041 Grenoble Cedex 9 France
2. Department of Botany and Zoology; Faculty of Science; Masaryk University; Kotlářská 2 Brno 61137 Czech Republic
3. Institute of Ecology and Evolutionary Biology; National Taiwan University; No. 1, Sec. 4, Roosevelt Rd. Taipei 10617 Taiwan
4. Dipartimento di Scienze; Università degli Studi di Roma Tre; V.le Marconi 446 - 00146 Roma Italy