1. Department of Animal Ecology; Netherlands Institute of Ecology (NIOO-KNAW); Droevendaalsesteeg 10 6708 PB Wageningen The Netherlands
2. Centre for Ecological and Evolutionary Synthesis (CEES); Department of Biosciences; University of Oslo; NO-0371 Oslo Norway
3. Division of Evolution, Ecology & Genetics; The Australian National University; Canberra ACT 2601 Australia