1. School of Life Sciences and Bioengineering, The Nelson Mandela African Institution of Science and Technology; Arusha 477 Tanzania
2. Department of Geography and Environmental Studies; University of Dodoma; Dodoma Tanzania
3. Department Conservation Biology; University of Dodoma; Dodoma Tanzania
4. Genome Science Centre and Department of Microbiology, Parasitology and Immunology; Sokoine University of Agriculture; Morogoro Tanzania
5. Center for Infectious Disease Dynamics, Huck Institutes of the Life Sciences and Department of Biology; Pennsylvania State University; University Park PA 16802 U.S.A.