1. RIKEN Center for Sustainable Resource Science, Tsurumi‐ku Yokohama Kanagawa 230‐0045 Japan
2. Division of Biological Science Nara Institute of Science and Technology Ikoma Nara 630‐0101 Japan
3. PMI R&D, Philip Morris Products S.A. Quai Jeanrenaud 5 CH‐2000 Neuchâtel Switzerland
4. Plant Molecular Science Center Chiba University, Chuo‐ku Chiba 260‐8675 Japan