1. Centre for Biodiversity and Environment Research, Department of Genetics, Evolution, and Environment; University College London; London WC1E 6BT United Kingdom
2. Department of Conservation Biology; Estación Biológica de Doñana (CSIC); Américo Vespucio 41092 Sevilla Spain
3. Department of Biology, Queens College; City University of New York; 65-30 Kissena Blvd Flushing New York NY 11367 USA
4. Ecology, Evolution and Behavior Subprogram, The Graduate Center; City University of New York; 365 5th Ave New York NY 10016 USA
5. Department of Zoology; Universidad de Sevilla; Avenida Reina Mercedes 6 41012 Sevilla Spain
6. Department of Physical, Chemical and Natural Systems; Universidad Pablo de Olavide; Ctra. Utrera km 1 41013 Sevilla Spain