1. Institute of Botany University of Basel 4056 Basel Switzerland
2. Research Unit Community Ecology WSL Swiss Federal Institute for Forest, Snow and Landscape Research Site Lausanne Station 2 CH‐1015 Lausanne Switzerland
3. Ecole Polytechnique Fédérale de Lausanne (EPFL) Laboratory of Ecological Systems (ECOS) School of Architecture, Civil and Environmental Engineering (ENAC) Station 2 CH‐1015 Lausanne Switzerland
4. WSL Swiss Federal Institute for Forest, Snow and Landscape Research 2000 Neuchâtel Switzerland
5. Institute of Geography University of Neuchâtel 2000 Neuchâtel Switzerland