1. Department of Ecology; Swedish University of Agricultural Sciences; PO Box 7044 SE-75007 Uppsala Sweden
2. Göteborg Museum of Natural History; PO Box 7283 402 35 Göteborg Sweden
3. Department of Ecology Science; Section Animal Ecology; Vrije Universiteit; 1081 HV Amsterdam The Netherlands
4. Groningen Institute for Evolutionary Life Sciences; Community and Conservation Ecology Group; University of Groningen; Nijenborgh 7 9747 AG Groningen The Netherlands