1. Key Laboratory of Algal Biology Institute of Hydrobiology Chinese Academy of Sciences Wuhan 430072 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Qingdao Branch of Shandong Water Diversion Project Operation and Maintenance Center Qingdao 266100 China
4. State Key Laboratory of Freshwater Ecology and Biotechnology Institute of Hydrobiology Chinese Academy of Sciences Wuhan 430072 China