1. Institute of Biology/Geobotany and Botanical Garden; Martin Luther University Halle-Wittenberg; Am Kirchtor 1 D-06108 Halle Germany
2. Herbario Nacional de Bolivia; Universidad Mayor de San Andrés; Correo Central, Casilla 10077 La Paz Bolivia
3. German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig; Deutscher Platz 5e D-04103 Leipzig Germany
4. Biodiversity and Climate Research Centre (BiK-F) and Senckenberg Gesellschaft für Naturforschung; Senckenberganlage 25 D-60325 Frankfurt (Main) Germany
5. Institute of Biological and Environmental Sciences; University of Aberdeen; St Machar Drive 23 AB24 3UU Aberdeen U.K