1. Key Laboratory of South China Agricultural Plant Molecular Analysis and Genetic Improvement Guangdong Provincial Key Laboratory of Applied Botany South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China
2. University of Chinese Academy of Sciences Chinese Academy of Sciences Beijing 100049 China
3. Guangdong Provincial Key Laboratory for Crop Germplasm Resources Preservation and Utilization Agricultural Biological Gene Research Center Guangdong Academy of Agricultural Sciences Guangzhou 510640 China
4. College of Food Science Southwest University Chongqing 400715 China
5. Institute of Plant Biology National Taiwan University Taipei 10617 Taiwan
6. Vegetable Research Institute Guangdong Academy of Agricultural Sciences Guangzhou 510640 China