1. USDA‐ARS, U.S. Pacific Basin Agricultural Research Center 64 Nowelo Street Hilo HI 96720 USA
2. Institute for Biodiversity and Ecosystem Dynamics (IBED) Evolutionary Biology and Population Biology University of Amsterdam Science Park 904 1098 XH Amsterdam The Netherlands
3. Université Côte d’Azur INRAE CNRS UMR ISA Nice France