1. Department of Integrative Biology, 501 Life Sciences West Oklahoma State University Stillwater Oklahoma 74078 USA
2. Mention Zoologie et Biodiversité Animale, Faculté des Sciences Université d'Antananarivo Antananarivo Madagascar
3. Museum of Natural Science and Department of Biological Sciences Lousiana State University Baton Rouge Louisiana 70803 USA
4. Department of Ecology and Evolutionary Biology University of Arizona Tucson Arizona 85721 USA