1. Zoological Researchmuseum Alexander Koenig; Adenauerallee 160 DE-53113 Bonn Germany
2. Dept of Biogeography; Trier Univ.; Universitätsring 15 DE-54286 Trier Germany
3. Dept of Biology, Molecular Ecology and Evolution Laboratory; Lund Univ., Ecology Building; SE-223 62 Lund Sweden
4. GECCO, Univ. of Angers; FR-49045 Angers France
5. UMR CNRS 6554 LETG Angers-LEESA, Univ. of Angers; FR-49045 Angers France
6. Dept of Animal and Plant Sciences; Univ. of Sheffield, Western Bank; Sheffield S10 2TN UK