1. The Key Laboratory of Zoological Systematics and Evolution Institute of Zoology Chinese Academy of Sciences Beijing China
2. Center for Excellence in Animal Evolution and Genetics Chinese Academy of Sciences Kunming China
3. Ecological Genetics Research Unit Research Programme in Organismal and Evolutionary Biology Faculty of Biological and Environmental Sciences University of Helsinki Helsinki Finland
4. Laboratory of Ichthyology Institute of Marine Biology Far East Branch of Russian Academy of Sciences Vladivostok Russia