1. Washington Cooperative Fish and Wildlife Research Unit; School of Aquatic and Fishery Sciences, University of Washington, Box 355020; Seattle, Washington, 98195 U.S.A.
2. U.S. Geological Survey, Washington Cooperative Fish and Wildlife Research Unit; School of Aquatic and Fishery Sciences, University of Washington, Box 355020; Seattle, Washington, 98195 U.S.A.