1. CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden Chinese Academy of Sciences Wuhan 430074 China
2. Center of Conservation Biology, Core Botanical Gardens Chinese Academy of Sciences Wuhan 430074 China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. School of Integrative Plant Science, Section of Plant Biology and the L.H. Bailey Hortorium Cornell University Ithaca NY 14850 USA
5. BTI Computational Biology Center, Boyce Thompson Institute Ithaca NY 14853 USA
6. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China