1. Key Laboratory of Aquatic Botany and Watershed Ecology, Wuhan Botanical Garden; Chinese Academy of Sciences; Wuhan 430074 China
2. Hubei Key Laboratory of Wetland Evolution & Ecological Restoration; Wuhan Botanical Garden, Chinese Academy of Sciences; Wuhan 430074 China
3. Donghu Experimental Station of Lake Ecosystems, State Key Laboratory of Freshwater Ecology and Biotechnology; Institute of Hydrobiology, Chinese Academy of Sciences; Wuhan 430072 China
4. Institute of Hydroecology; Ministry of Water Resources & Chinese Academy of Sciences; Wuhan 430079 China
5. Faculty of Agriculture, Laboratory of Plant Conservation Science; Meijo University; Aichi 468-8502 Japan