1. Key Laboratory for Plant Diversity and Biogeography of East Asia Kunming Institute of Botany, Chinese Academy of Sciences Kunming 650201 China
2. Germplasm Bank of Wild Species Kunming Institute of Botany, Chinese Academy of Sciences Kunming 650201 China
3. Department of Phytopathology, College of Plant Protection Yunnan Agriculture University 650201 China
4. Royal Botanic Gardens, Kew, Wakehurst Place West Sussex RH17 6TN UK