1. Food Crops Institute, Hubei Academy of Agricultural Sciences, Hubei Key Laboratory of Food Crop Germplasm and Genetic Improvement Laboratory of Crop Molecular Breeding, Ministry of Agriculture and Rural Affairs Wuhan China
2. National Key Laboratory of Crop Genetic Improvement and National Center of Plant Gene Research, College of Life Science and Technology Huazhong Agricultural University Wuhan China
3. Hubei Hongshan Laboratory Wuhan China