1. School of Natural Sciences, College of Environmental Sciences and Engineering, Bangor University Bangor UK
2. Biodiversity Inventory for Conservation NPO (BINCO) Glabbeek Belgium
3. Wild Planet Trust Paignton Zoo Paignton UK
4. Department of Biological Sciences Simon Fraser University Burnaby British Columbia Canada
5. Terrestrial Ecology Research Group, Department for Life Science Systems, School of Life Science Technical University of Munich Freising Germany