1. Key Laboratory of Biogeography and Bioresource in Arid Land; Xinjiang Institute of Ecology and Geography; Chinese Academy of Sciences; Urümqi China
2. Turpan Eremophytes Botanical Garden; Chinese Academy of Sciences; Turpan China
3. University of the Chinese Academy of Sciences; Beijing China
4. Department of Biology; University of Kentucky; Lexington Kentucky USA
5. Department of Plant and Soil Sciences; University of Kentucky; Lexington Kentucky USA