1. Graduate School of Agriculture Kyoto University Kyoto Japan
2. Graduate School of Life Sciences Tohoku University Sendai Miyagi Japan
3. Koishikawa Botanical Garden, Graduate School of Science University of Tokyo Tokyo Japan
4. Tropical Biosphere Research Center University of the Ryukyus Taketomi Okinawa Japan
5. Faculty of Regional Environment Science Tokyo University of Agriculture Tokyo Japan
6. Biological Resources and Post‐harvest Division Japan International Research Center for Agricultural Sciences Tsukuba Ibaraki Japan